C. pictipennis (Staeger 1839)

Taxonomy	Biology	Distribution

Ceratopogon pictipennis Staeger 1839: 594. TL Denmark
(Kieffer 1919: 46. ♀, note)
(Edwards 1926: 406. Notes)
(Kettle & Lawson 1952: 439. Early stages, key)
(Campbell & Pelham-Clinton 1960: 243. ♂, ♀, notes, keys, figs)
(Gutsevich 1960: 81. ♂, ♀, notes, figs)
(Kremer 1966: 138. ♂, ♀, notes, figs)
(Navai & Mesghali 1968: 244. Record, Iran)
(Kremer, Callot & Hommel 1973: 61. Notes, figs, key)
(Glukhova 1979: 201. Larva)
(Remm 1981: 29. Synon)
(Delecolle 1985: 183. ♂, ♀, notes, figs)
(Boorman 1986: 262. Notes)
(Glukhova 1989: 234. ♂, ♀, notes, figs)
    = arcuatus (Winnertz)
Ceratopogon arcuatus Winnertz 1852: 39. TL Germany
(Goetghebuer 1920: 53. Note)
(Edwards 1926: 406. Notes)
    = guttularis Kieffer
Culicoides guttularis Kieffer 1919: 45. TL Hungary
(Kieffer 1919: 45. ♀, desc, fig)
(Austen 1921: 114. Note)
(Austen 1925: 2. note, record, Dardanelles)
    = maculatus Zilahi-Sebess
Culicoides pictipennis var. maculatus Zilahi-Sebess 1936: 202. TL Hungary (preocc.)
    = subgrisescens Dzhafarov
Culicoides subgrisescens Dzhafarov 1964: 249. TL Azerbaijan
(Glukhova 1989: 239. ♂, ♀, notes, figs)
    = achkamalicus Dzhafarov
Culicoides achkamalicus Dzhafarov 1964: 251. TL Azerbaijan
(Kremer, Callot & Hommel 1973: 61. Notes, figs, key)
    = luganicus Shevchenko
Culicoides luganicus Shevchenko 1972: 75. TL Ukraine

Diagnosis and Notes

The wings are greyish with prominent rounded pale spots; the second radial cell wholly within the second costal spot, and the pale spots at the tips of cells r5, m1 and m2 are cut by the wing margin. AR 1.64-1.73; SD 3-15. In the male, the ninth sternite membrane is bare. It is very like griseidorsum, which has antennal sensilla on 3-14; and also like saevanicus, in which the two spots in the anal cell are only narrowly separated by a darker band. It differs from univittatus in that the marginal pale spots are cut by the wing margin. It could also easily be mistaken for poperinghensis, but this has the second radial cell only partly pale, the wing spots less well defined, and the female SD 3,11-15.
The figure of the wing is from Edwards (1939) and of the male genitalia from Glukhova (1989).