C. punctatus (Meigen 1804)

Taxonomy	Biology	Distribution

Ceratopogon punctatus Meigen 1804: 29. TL Europe
(Edwards 1939: 47. Notes, figs, as pulicaris var.)
(Downes & Kettle 1952: 70. Notes, figs, key)
(Kettle & Lawson 1952: 445. Early stages, key)
(Campbell & Pelham-Clinton 1960: 277. ♂, ♀, notes, keys, figs)
(Coluzzi & Kremer 1964: 67. Notes, record, Italy)
(Callot, Kremer & Coluzzi 1965: 162. Notes, Italy
(Smatov 1966: 83. Notes, record Kazakhstan)
(Kremer 1966: 199. ♂, ♀, notes, figs)
(Navai & Mesghali 1968: 245. Record, Iran)
(Remm 1973: 174. ♂, ♀, record, Mongolia)
(Havelka 1976: 224. Notes)
(Glukhova 1979: 190. Larva)
(Havelka 1982: 85. ♂, ♀, notes, figs)
(Boorman 1986: 257. Notes)
(Delecolle 1985: 24. ♂, ♀, notes, figs)
(Glukhova 1989: 162. ♂, ♀, notes, figs)
    = punctatus Latreille
Ceratopogon punctatus Latreille 1809: 252 TL France (preocc)
    = ocellaris Kieffer
Culicoides pulicaris var. ocellaris Kieffer 1921: 276. TL Latvia
    = kasachstanicus Shakirzjanova
Culicoides pulicaris ssp. kasachstanicus Shakirzjanova 1963: 63. TL Kazakhstan
(Gutsevich & Smatov 1966: 70. ♂, ♀, notes, figs)
(Smatov 1966: 83. Notes, record Kazakhstan)
    = remmi Damian-Georgescu
Culicoides remmi Damian-Georgescu 1972: 16. TL Rumania

Diagnosis and Notes

The tips of wing veins M1, M2 and CU1 typically end at the wing tip in small pale spots, distinguishing this species from C. pulicaris; but the distinction is not always clear-cut. Both of these species are variable in the extent and intensity of the wing markings (see also Lane (1981)). The female AR is 1.02-1.15; the SD 3,11-15. The figure of the wing is from Edwards (1939), and of the male genitalia from Kremer (1966).